Bird()
constructor.computeFlightPerformance()
.A bird can be defined using the Bird()
construction
function. This function ensures that the generated object contains all
required parameters that describe the bird.
myBird <- Bird(
massTotal = 0.215, # weight in kg
wingSpan = 0.67, # wingspan in m
wingArea = 0.0652, # wing area in m2
name = 'Jackdaw',
name.scientific = 'Corvus monedula',
type = 'passerine',
source = 'KleinHeerenbrink M, Warfvinge K and Hedenstrom A (2016) J.Exp.Biol. 219: 10, 1572--1581'
)
Any parameters that are not specified, but that are required by any
of the flight performance calculations will be filled in with either
defaults or estimated with allometric scaling relationships. For
instance, the body frontal area, that is used for computing body drag,
is in this case calculated based on the specified massTotal
from an allometric relationship for type='passerine'
:
## [1] 0.005020037
It is possible to modify the bird description simply by reassigning the variables in the list, i.e.
However, it should be noted that some of the variables are related to
others (e.g. massTotal
is the sum of
massEmpty
, massFat
and massLoad
),
so that manual changes can lead to unexpected results.
The default bird settings mostly follow the suggestions in Pennycuick (2008).
Flight performance can be computed using
computeFlightPerformance()
. It calculates maximum available
mechanical power from the muscles, aerodynamic (mechanical) power for
flapping flight as a function of airspeed and the corresponding chemical
power.
## Name: Jackdaw
## Sc. name: Corvus monedula
## Bird definitions: KleinHeerenbrink M, Warfvinge K and Hedenstrom A (2016) J.Exp.Biol. 219: 10, 1572--1581
## speed power.aero power.chem strokeplane amplitude
## minimumSpeed 3.398 4.532 23.93 49.9 53.6
## minimumPower 8.144 2.167 12.62 25.3 37.6
## maximumRange 11.110 2.603 14.71 16.9 40.2
## maximumSpeed 15.433 4.531 23.93 10.6 51.2
## Maximum climb performance:
## speed power.aero power.chem strokeplane amplitude climbRate
## maximumClimbRate 8.886 4.531 23.93 22.6 53.5 0.8587
## Minimized migration time:
## speed speed.migration power.aero power.chem power.dep
## minimumTimeSpeed 11.82 1.934 2.819 15.74 3.081
## strokeplane amplitude
## minimumTimeSpeed 15.55 41.62
This is the simplest call to computeFlightPerformance()
.
The print()
of the flightperf
shows a summary
of the computed powercurve. The function
computeFlightPerformance()
has calculated several
characteristic airspeeds:
minimumSpeed
: the required aerodynamic power equals the
maximum available muscle power1.minimumPower
: the required aerodynamic/chemical power
is minimal, i.e. this is the speed for maximum endurance.maximumRange
: the cost of transport is minimal, i.e
this is the speed for maximum range.maximumSpeed
: required aerodynamic power equals the
maximum available muscle power.These characteristic speeds are stored as a data.frame
named table
, which contains several more advanced variables
not shown in the print
, as explained in the following
sections.
The function computeFlightPerformance()
also estimates
the best climb performance, i.e. the maximum climb rate that can be
achieved given the power available from the muscles. This data point is
stored in the data.frame
named maxClimb
. It
contains the same variables as table
, plus an additional
one: climbRate
. Similarly, the function also produces a
data.frame
named minTime
, which holds the
flight performance for time minimizing migratory flight. At this
airspeed, which will be slightly higher than the maximum range speed,
the bird balances flight time with the time it takes to (re)gain the
energy lost during flight.
Finally, computeFlightPerformance()
also computes a
powercurve for a range of speeds. The length of this curve can be
specified by providing a length.out
argument, but the
default produces 10 points. This curve can be used to visualize the
speed dependency of the various variables.
powercurve <- flightperf$powercurve
plot(flightperf,symbol.speed="U")
plot(powercurve$speed,powercurve$amplitude,xlab=NA,ylab=NA,type='b',xlim=c(0,20),ylim=c(20,60))
mtext(side = 1, line = 2, 'Airspeed (m/s)')
mtext(side = 2, line = 2, 'Peak amplitude (deg)')
In the current example, the strokeplane is optimized. The model also implicitly optimizes the wingbeat amplitude for minimum induced power, so essentially the returned powercurve represents the optimal wingbeat for a given wingbeat frequency.
## speed frequency strokeplane amplitude
## minimumSpeed 3.397602 5 49.94069 53.63391
## minimumPower 8.144311 5 25.30698 37.55072
## maximumRange 11.109520 5 16.89529 40.18648
## maximumSpeed 15.433460 5 10.60542 51.16479
By default, computeFlightPerformance()
will try to
optimize the strokeplane angle for minimum power at any given speed.
This optimization is performed many times, which results in a high
computational load. The optimization can be avoided by explicitly
specifying a strokeplane angle,
e.g. computeFlightPerformance(myBird,strokeplane=0)
.
table
contains flags that indicate whether the data
point is outside the validity range of the model:
## flags.redFreqLo flags.redFreqHi flags.thrustHi flags.speedLo
## minimumSpeed FALSE TRUE TRUE FALSE
## minimumPower FALSE FALSE FALSE FALSE
## maximumRange FALSE FALSE FALSE FALSE
## maximumSpeed FALSE FALSE FALSE FALSE
In this case the flags indicate the the minimum speed is outside the
model’s validity range, indicating too high reduced frequency and too
high thrust requirement. The model was obtained for reduced frequencies
\(1 \leq k_f \leq 6\) and thrust
requirements (thrust to lift ratio) \(0 \leq
\frac{T}{L} \leq 0.3\). The flag flags.speedLo
indicates that the data point is below the low speed limit of the model
(\(U<2w_\mathrm{i,h}\); airspeed is
lower than twice the induced downwash in hover2). Data points with
flags may be reasonable extrapolations of the model, but it is safer to
assume flagged data points are invalid.
Stored in the table
are also the drag components. Only
the non-flapping (Dnf
) components are stored:
## Dnf.ind Dnf.pro0 Dnf.pro2 Dnf.par
## minimumSpeed 0.44582824 0.008148623 0.28929485 0.007098852
## minimumPower 0.07758968 0.030241749 0.05034741 0.040789816
## maximumRange 0.04169866 0.048180164 0.02705797 0.075898565
## maximumSpeed 0.02160658 0.078889703 0.01402036 0.146477065
These are the induced drag, zero lift profile drag, lift dependent profile drag and parasitic drag, respectively. The latter includes the body drag, but also any other additional drag (e.g. the drag component of weight during climb).
par(mar=c(3.1,3.1,0.4,1.1),mgp=c(1.9,.7,0),cex=0.75)
with(powercurve , plot( speed, Dnf.ind, type='b', col='red3',
xlab=NA, ylab=NA, xlim=c(0,20), ylim=c(0.0,0.20)))
with(powercurve , lines( speed, Dnf.pro0, type='b', col='green3'))
with(powercurve , lines( speed, Dnf.pro2, type='b', col='blue3'))
with(powercurve , lines( speed, Dnf.par, type='b', col='yellow3'))
mtext(side = 1, line = 2,'Airspeed (m/s)')
mtext(side = 2, line = 2,'Drag (N)')
The above figure shows the magnitude of the non-flapping drag components for different speeds. The lift dependent components are very large at low speeds and they decrease rapidly with increasing speed. The parasitic drag shows the classic quadratic behaviour with speed. The zero-lift profile drag is near zero at low speeds, and increases more linearly with speed. This is due to the strong dependence of the friction coefficient on the Reynolds number. As non-flapping effectively means gliding, these drag components can be used to construct a drag polar for gliding flight. However, this ignores the wing flex that typically occurs at higher speeds.
Flapping the wings alters the drag experienced by the wings: \(D = k_D D^\prime\) (using \(D^\prime\) for non-flapping drag) 3. The
factors \(k_D\) depend on the wingbeat
kinematics and are returned in table
:
## kD.ind kD.pro0 kD.pro2
## minimumSpeed 0.942585 2.4759503 0.7855803
## minimumPower 1.327851 1.0184287 1.2634913
## maximumRange 1.466031 0.9736113 1.4052916
## maximumSpeed 1.875661 0.9167451 1.7971466
These factors are for induced drag, zero lift profile drag and lift dependent profile drag respectively.
The aerodynamic power components for flapping flight can be computed
in a similar way: \(P = k_P D^\prime
U\). The factors \(k_P\) depend
on the wingbeat kinematics and are also stored in
table
:
## kP.ind kP.pro0 kP.pro2
## minimumSpeed 1.928459 2.8711163 1.533473
## minimumPower 1.554799 1.0547097 1.444999
## maximumRange 1.636867 0.9964092 1.557558
## maximumSpeed 2.045005 0.9506620 1.990849
par(mar=c(3.1,3.1,0.4,1.1),mgp=c(1.9,.7,0),cex=0.75)
with(powercurve , plot( speed, kD.ind, type='b', col='red3',
xlab=NA, ylab=NA, xlim=c(0,20), ylim=c(0.5,2.5)))
with(powercurve , lines( speed, kD.pro0, type='b', col='green3'))
with(powercurve , lines( speed, kD.pro2, type='b', col='blue3'))
mtext(side = 1, line = 2,'Airspeed (m/s)')
mtext(side = 2, line = 2,'Drag factors (-)')
with(powercurve , plot( speed, kP.ind, type='b', col='red3',
xlab=NA, ylab=NA, xlim=c(0,20), ylim=c(0.5,2.5)))
with(powercurve , lines( speed, kP.pro0, type='b', col='green3'))
with(powercurve , lines( speed, kP.pro2, type='b', col='blue3'))
mtext(side = 1, line = 2,'Airspeed (m/s)')
mtext(side = 2, line = 2,'Power factors (-)')
The above figure shows the speed dependency of the drag factors and power factors. At low speeds the lift-dependent drag factors have a low value. There is even a tendency for the factors to drop below 1. The zero-lift profile drag instead takes very large values. At these low speeds the velocity of the wings starts to become more dominant than the flight speed.
Several other variables are also stored in table
:
## frequency L ReynoldsNumber CDpro0
## minimumSpeed 5 2.10843 22646.05 0.017676057
## minimumPower 5 2.10843 54284.31 0.011416802
## maximumRange 5 2.10843 74048.32 0.009775163
## maximumSpeed 5 2.10843 102868.70 0.008293541
These are respectively the wingbeat frequency (in this case as
specified in myBird
), the lift force (in this case just
equal to weight), the mean chord Reynolds number, and the zero lift
profile drag coefficient (dependent on mean chord Reynolds number).
All models by default use the internal FLIGHTCONDITION
,
which describes the standard atmosphere, with air density \(\rho=1.225\) kg m3,
gravitational acceleration \(g=9.81\) m
s-2 and kinematic viscosity \(\nu=14.61\times 10^{-6}\) m2
s-1. It is possible to define an alternative flight
condition:
myFlightCondition <- list(
density = 0.9093, # [kg/m3] 3 km altitude
gravity = 9.81, # [m/s2]
viscosity = 18.63E-6, # [m2/s] 3 km altitude
windSpeed = 0,# [m/s]
windDir = 0 # [degrees] 0 degrees means wind is along the track direction
)
Air density decreases with altitude, and a density of \(\rho=0.909\) kg m3 corresponds to a flight altitude of 3 km above sea level in the international standard atmosphere. At this altitude the kinematic viscosity has increased to \(\nu=18.63\times 10^{-6}\) m2 s-1. This alternative flight condition can be used in the performance calculation as:
flightperf.ISA3 <- computeFlightPerformance(myBird,flightcondition=myFlightCondition)
flightperf.ISA3
## Name: Jackdaw
## Sc. name: Corvus monedula
## Bird definitions: KleinHeerenbrink M, Warfvinge K and Hedenstrom A (2016) J.Exp.Biol. 219: 10, 1572--1581
## speed power.aero power.chem strokeplane amplitude
## minimumSpeed 4.946 4.531 23.93 46.2 54.4
## minimumPower 9.556 2.634 14.85 22.8 40.7
## maximumRange 12.873 3.129 17.22 15.6 43.5
## maximumSpeed 16.372 4.531 23.93 11.5 51.1
## Maximum climb performance:
## speed power.aero power.chem strokeplane amplitude climbRate
## maximumClimbRate 8.886 4.531 23.93 22.6 53.5 0.8587
## Minimized migration time:
## speed speed.migration power.aero power.chem power.dep
## minimumTimeSpeed 13.57 1.961 3.339 18.23 3.081
## strokeplane amplitude
## minimumTimeSpeed 14.56 44.79
Note how the power requirements have increased (this is only apparent
from the minimumPower
and maximumRange
, as the
maximum available power is unchanged), but the characteristic speeds
also increased. Looking at the energetic cost per traveled distance:
COT.ISA0 <- with(flightperf$table,power.chem/speed)
COT.ISA3 <-with(flightperf.ISA3$table,power.chem/speed)
list(ISA0 = COT.ISA0, ISA3 = COT.ISA3)
## $ISA0
## [1] 7.044379 1.549928 1.323949 1.550427
##
## $ISA3
## [1] 4.838110 1.554379 1.337836 1.461550
## [1] 0.6868043 1.0028719 1.0104890 0.9426758
it appears that flying at the maximum range speed at a higher altitude will actually reduce the maximum range (which can be attributed to the increase in kinematic viscosity, increasing the profile drag coefficient). However, the 1% increase in flight cost does pay back as a 15% reduction in flight time. In fact, if the bird cares about time minimization, flying at altitude is highly advantageous, increasing the maximum flight speed while reducing the associated cost of transport.